Report on genital studies of European Clausiliidae for taxonomic and phylogenetic work
Hartmut Nordsieck (XII. 2020, updated (not pdf) IX.2021)
I. General remarks, Material and methods
Like in other stylommatophoran families the genitalia of the species of Clausiliidae,especially the copulatory organs, are of high taxonomic importance and thus of interest for phylogenetic studies. Therefore, for a book on European Clausiliidae I studied the genitalia of about 330 species of all subfamilies by describing and measuring about 550 preparations, which I had made from alcohol material of these species (1-2, in part more specimens prepared). The material used and the preparations are stored in the Staatliches Museum für Naturkunde Stuttgart (SMNS) and the Forschungsinstitut Senckenberg Frankfurt am Main (SMF).
In this article my former papers on genital morphology are cited by the publication years.
The material comprises the following genera (in brackets number of examined species):
Pontophaedusa (1), Pontophaedusella (1), Laeviphaedusa (1),
Caspiophaedusa (1), Pravispira (1), Dobatia (1), Serrulina (2).
For the tribe system see article on DNA studies.
Alopiini (sensu lato):
Carinigera (16), Isabellaria (8), Albinaria (30), Inchoatia (3), Cristataria (6),
Medora (7), Agathylla (8), Lampedusa (1), Muticaria (1), Leucostigma (1),
Strigilodelima (1), Montenegrina (7), Protoherilla (1), Herilla (7), Alopia (20).
Charpentieria (5), Siciliaria (17), Papillifera (2), Delima (16), Dilataria (3), Barcania (1).
Former Mentissoideinae + Graciliaria:
Boettgeria (6), Euxinella (1), Olympicola (1), Filosa (1), Scrobifera (1),
Strigileuxina (2), Sumelia (2), Akramowskia (1), Acrotoma (2),
Roseniella (2), Armenica (5), Inobseratella (2), Phrygica (2), Sprattia (3),
Euxina (8), Elia (7), Euxinastra (2), Galeata (3), Mentissoidea (1),
Idyla (3), Graciliaria (1), Micridyla (1).
Ruthenica (1), Pseudofusulus (1), Erjavecia (1),
Julica (1), Macrogastra (10), Clausilia (10), Fusulus (2).
Quadriplicata (2), Mucronaria (3), Micropontica (1), Mentissa (1), Mentissella (1),
Laciniaria (3), Alinda (4), Pseudalinda (4), Strigillaria (12), Vestia (4), Pavlovicia (2), Balea (4).
For definition and terms of the parts of the copulatory organs see 2007a (: chapter I: 9-11, 16, appendix 2: 178-180). The terms proximal and distal are used as seen from the gonad.
The measurements were made as exactly as possible making use of an ocular micrometer (accuracy ± 0.1 mm). Before measuring the preparations were stretched as far as possible without tearing and fixed by micropins without piercing. It is important for measurements to know about the limitation of the parts (see Fig. 1). The limitation of the female parts are given by the branchings. The limitation of the male parts are more difficult to make out, especially in groups in which they are more or less modified (former Serrulininae, Clausiliini, Baleini). They are located by changes of the lumen and the wall structure.