Vietnamese Clausiliidae

Vietnamese Clausiliidae

List of Vietnamese Clausiliidae

Hartmut Nordsieck (II.2015)

The species are listed in alphabetical order; assumed subspecies and synonyms (indicated with =) are added. The species are accompanied by information on their known geographical range (provinces).
The list, which for the first time was published in Nordsieck (2011: 150-152), is completed by the collecting results of C. & J. Hemmen in 2011 and 2012 and the information given by Grego et al. (2014).

Phaedusinae A. J. Wagner 1922

abletti Pham & Szekeres 2014, Phaedusa (P.): Dien Bien.
abstrusa (Szekeres 1970), Oospira (O.) (aabstrusaaginkae Grego & Szekeres 2014): Dien Bien, Hoa Binh, Ninh Binh, Thanh Hoa.
aciculata (Bavay & Dautzenberg 1909), Oospira (Lindholmiella): Lai Chau.
acrostoma (Bavay & Dautzenberg 1909), Synprosphyma (S.): Lao Cai, Lai Chau.
antibouddah H. Nordsieck 2003, Oospira (Atractophaedusa): Quang Ninh.
auregani (Bavay & Dautzenberg 1903), Liparophaedusa: Lang Son, Bac Can, Thai Nguyen.
auricoma (Bavay & Dautzenberg 1899), Synprosphyma (S.): Bac Can.
babeensis (Bavay & Dautzenberg 1899), Synprosphyma (S.) (bbabeensisbcrassitesta H. Nordsieck 2011): Bac Can, Thai Nguyen.
bavayi H. Nordsieck 2002, Hemiphaedusa (Selenophaedusa): Lao Cai, Bac Can, Lang Son.
billeti (Fischer 1898), Hemiphaedusa (Selenophaedusa) (= falcifera Bavay & Dautzenberg 1899, = callistomella Bavay & Dautzenberg 1900) (bbilleti): Ha Giang, Cao Bang, Lang Son, Bac Can.
bouddah (Bavay & Dautzenberg 1912), Oospira (O.): Quang Ninh, Ha Noi, Hoa Binh.
callistoma (Bavay & Dautzenberg 1899), Hemiphaedusa (Selenophaedusa):Cao Bang, Lang Son, Bac Can.
cervicalis (Bavay & Dautzenberg 1909), Synprosphyma (S.): Lao Cai.
chiemhoaensis (Sykes 1902), Hemiphaedusa (Selenophaedusa) (= lavillei Dautzenberg & Fischer 1905): Ha Giang, Tuyen Quang.
cochinchinensis (L. Pfeiffer 1841), Phaedusa (P.): Da Nang.
coudeini (Bavay & Dautzenberg 1899), Oospira (Formosana): Lang Son, Bac Can.
cuongi Maassen & Gittenberger 2007, Oospira (O.) (ccuongic. n. subsp. H. Nordsieck): Ha Noi, Hoa Binh, Thanh Hoa.
dextrogyra (Bavay & Dautzenberg 1909), Oospira (Formosana): Lao Cai.
dichroa (Bavay & Dautzenberg 1899), Phaedusa (P.) (ddichroadduporti Bavay & Dautzenberg 1915): Lang Son, Bac Can, Ninh Binh, Hoa Binh.
diplochilus (Moellendorff 1901), Hemiphaedusa (Selenophaedusa): Lang Son.
duci Maassen & Gittenberger 2007, Oospira (O.) (dducidkhanhi H. Nordsieck 2011, d. n. subsp. 1 H. Nordsieck 1, d. n. subsp. 2 H. Nordsieck): Son La, Ha Noi, Thanh Hoa, Quang Binh.
duella (Mabille 1887), Dautzenbergiella (D.) (dapiostoma Moellendorff 1901 (= lyteostoma Moellendorff 1901), dduella): Lang Son.
eregia (Szekeres 1969), Oospira (O.): Ninh Binh, Nghe An.
fistulata (Bavay & Dautzenberg 1909), Synprosphyma (S.): Lao Cai, Ha Giang, Tuyen Quang.
freyi (Bavay & Dautzenberg 1899), Liparophaedusa: Bac Can, Thai Nguyen.
gastrodes (Moellendorff 1901), Synprosphyma (S.): Lang Son.
gastrum (Moellendorff 1901), Synprosphyma (S.) (= ventriculus Moellendorff 1901): Lang Son.
grangeri (Bavay & Dautzenberg 1899), Oospira (Siphonophaedusa) (ggrangeri): Cao Bang, Lang Son.
kebavica (Moellendorff 1901), Oospira (Atractophaedusa): Quang Ninh.
lemyrei (Bavay & Dautzenberg 1899), Hemiphaedusa (Dendrophaedusa): Lang Son, Bac Can, Thai Nguyen.
leptotesta H. Nordsieck 2011, Liparophaedusa: Tuyen Quang.
lypra (Mabille 1887), Phaedusa (P.) (leupleura Bavay & Dautzenberg 1899, linanis Bavay & Dautzenberg 1909, llypra ( = houssayi Fischer 1898, = backanensis Bavay & Dautzenberg 1899, = hamonvillei Bavay & Dautzenberg 1899), l. n. subsp. H. Nordsieck, lpereupleura H. Nordsieck 2011, lsubgranulosa H. Nordsieck 2003): Cao Bang, Lang Son, Bac Can, Thai Nguyen, Vinh Phuc, Ninh Binh, Hoa Binh, Son La, Lao Cai, Lai Chau, Dien Bien, Thanh Hoa, Nghe An.
mairei (Bavay & Dautzenberg 1909), Oospira (O.): Lao Cai, Lai Chau.
mansuyi (Dautzenberg & Fischer 1908), Dautzenbergiella (Mansuyiella) (mmansuyi): Cao Bang.
margaritifera (Bavay & Dautzenberg 1909), Hemiphaedusa (Margaritiphaedusa): Lao Cai.
micropaviei H. Nordsieck 2011, Phaedusa (P.): Son La.
mirabilis H. Nordsieck 2003, Thaumatoptyx?: Lao Cai.
miranda (Loosjes & Loosjes-van Bemmel 1973), Oospira (O.): Ninh Binh, Hoa Binh, Thanh Hoa, Nghe An.
moirati (Bavay & Dautzenberg 1909), Synprosphyma (S.) (= rudis A. J. Wagner 1922): Lao Cai, Lai Chau.
n. sp. H. Nordsieck, Oospira (O.): Son La.
n. sp. H. Nordsieck, Synprosphyma (S.): Son La.
naggsi Luong & Szekeres 2014, Oospira (O.): Dong Nai.
oospiroides H. Nordsieck 2011, Synprosphyma (S.): Ha Giang.
ophthalmophana (Mabille 1887), Hemiphaedusa (Selenophaedusa) (ocazioti Bavay & Dautzenberg 1909, oophthalmophana): Lang Son, Thai Nguyen, Hoa Binh.
oviformis H. Nordsieck 2011, Oospira (O.): Son La, Hoa Binh, Thanh Hoa.
paviei (Morlet 1892), Phaedusa (P.): Lai Chau, Lao Cai, Son La, Ha Giang, Tuyen Quang, Bac Can, Lang Son, Yen Bai, Vinh Phuc, Ha Noi.
phongthoensis Loosjes & Loosjes-van Bemmel 1949, Phaedusa (P.): Lai Chau.
polydona (Mabille 1887), Hemiphaedusa (Notoptychia) (pmontana Moellendorff 1901, ppolydona (= gisota Bavay & Dautzenberg 1899, = platyloma Moellendorff 1901)): Lang Son, Bac Can.
porphyrea (Moellendorff 1882), Euphaedusa (Papilliphaedusa) (ptheristica Mabille 1887 (= vatheleti Bavay & Dautzenberg 1899, = gereti Bavay & Dautzenberg 1900): Ha Giang, Cao Bang, Lang Son, Bac Can, Tuyen Quang, Thai Nguyen, Ha Noi, Hoa Binh, Son La.
porphyrostoma (Bavay & Dautzenberg 1909), Hemiphaedusa (Selenophaedusa) (pporphyrostomapregina H. Nordsieck 2011): Lao Cai, Ha Giang.
pseudauregani (Dautzenberg & Fischer 1908), Liparophaedusa: Cao Bang.
pumilio H. Nordsieck 2003, Hemiphaedusa (Hemizaptyx): Lao Cai.
pyknosoma Gittenberger & Vermeulen 2001, Oospira (Atractophaedusa): Hai Phong.
rhopaloides (Moellendorff 1901), Oospira (Atractophaedusa) (= leptospira Moellendorff 1901, = microthyra Moellendorff 1901): Quang Ninh.
ruminiformis (Mabille 1887), Liparophaedusa: Lang Son.
semipolita (Bavay & Dautzenberg 1899), Oospira (O.) (smansonensis Moellendorff 1901, ssemipolita (= flaveola Bavay & Dautzenberg 1915)): Lao Cai, Ha Giang, Tuyen Quang, Bac Can, Lang Son.
smithi Maassen & Gittenberger 2007, Oospira (Atractophaedusa): Hai Phong.
sorella H. Nordsieck 2003, Phaedusa (P.): Lao Cai, Bac Can.
stenothyra Moellendorff 1901, Phaedusa (P.): Quang Ninh, Hai Phong, Ninh Binh, Hoa Binh, Thanh Hoa.
suilla (Bavay & Dautzenberg 1909), Synprosphyma (S.): Lao Cai, Lai Chau.
sykesi (Bavay & Dautzenberg 1899), Oospira (Leptocochlea) (sletrungthongi Grego & Szekeres 2011, ssykesi): Lang Son, Bac Can, Thai Nguyen.
thatkheana (Bavay & Dautzenberg 1899), Hemiphaedusa (Selenophaedusa) (tsplendida H. Nordsieck 2011, tthatkheana): Ha Giang, Cao Bang, Lang Son, Bac Can, Tuyen Quang.
triptyx H. Nordsieck 2011, Oospira (O.): Son La.
vanbuensis (Bavay & Dautzenberg 1899), Oospira (O.) (= pocsi Szekeres 1969, = umbratica Szekeres 1970): Ha Noi, Son La, Hoa Binh, Ninh Binh, Thanh Hoa, Nghe An.

Garnieriinae C. Boettger 1926

The synonymization of Neniauchenia and Grandinenia by Grego et al. (2014) is not accepted. The genital apparatus of Neniauchenia steffeki (: fig. 1) is, as far as can be concluded from the figure, largely correspondent with that of Ptychauchenia panhai (Nordsieck 2010: text-fig. 1). Both differ from that of all (six) examined Grandinenia species by the shorter penis, especially its proximal part, and the longer epiphallus which is bent before the transition to the penis. Neniauchenia and Ptychauchenia exhibit also a lower palatal plica, which is quite different from that of Grandinenia (well-developed, plica-like). Neniauchenia is obviously more closely related to Ptychauchenia than to Grandinenia. A combination of Neniauchenia and Grandinenia within one genus is therefore not recommendable.

ardouiniana (Heude 1885), Grandinenia (= horrida Mabille 1887, = sajoi Szekeres 1969): Quang Ninh, Hai Phong.
bavayi (Lindholm 1924), Tropidauchenia (bbavayibsenescens H. Nordsieck 2011): Bac Can, Tuyen Quang.
demangei (Bavay & Dautzenberg 1909), Euryauchenia (ddemangeiddextroversa H. Nordsieck 2014): Ninh Binh, Thanh Hoa.
dorri (Bavay & Dautzenberg 1899), Tropidauchenia: Bac Can.
fischeri H. Nordsieck 2002, Euryauchenia (ffischerifreticulata H. Nordsieck 2002 (= phasmoides Grego & Szekeres 2011)): Tuyen Quang?, Ninh Binh, Hoa Binh, Thanh Hoa.
gabijakabi (Grego & Szekeres 2014), Neniauchenia: Quang Binh.
giardi (Fischer 1898), Tropidauchenia (gbrunnea H. Nordsieck 2002, gcitrina H. Nordsieck 2002, ggiardi): Cao Bang, Lang Son, Bac Can.
ootanii H. Nordsieck 2007, Tropidauchenia (olongicollis H. Nordsieck 2011): Ha Giang.
orientalis (Mabille 1887), Tropidauchenia (omessageri Bavay & Dautzenberg 1899, oorientalis): Ha Noi, Lang Son.
palatalis H. Nordsieck 2011, Tropidauchenia: Ha Giang.
panhai H. Nordsieck 2010, Ptychauchenia (p. n. subsp. H. Nordsieck, ppanhai): Quang Binh.
proctostoma (Mabille 1889), Megalauchenia (pdonggiaoensis (H. Nordsieck 2002), pforceps (Loosjes & Loosjes-van Bemmel 1973), pproctostoma (= goliath Rolle 1910)): Ha Nam, Ninh Binh, Hoa Binh, Son La, Thanh Hoa.
soror H. Nordsieck 2011, Megalauchenia: Thanh Hoa.
steffeki (Grego & Szekeres 2014), Neniauchenia: Nghe An.
tonkinensis H. Nordsieck 2010, Neniauchenia: Quang Binh.
yongshuae H. Nordsieck 2007, Megalauchenia: Thanh Hoa.

According to the information of S. Panha on the respective labels, Ptychauchenia panhai (Fig. 1) and Neniauchenia tonkinensis (Fig. 2) have the type locality Cuc Phuong N. P., Ninh Binh. Both species have otherwise only been collected in the province Quang Binh; neither by K. Ookubo nor by C. & J. Hemmen have they been found in Cuc Phuong N. P.
Checking the Senckenberg collection I stated that also Megalauchenia proctostoma and Oospira miranda have been recorded on labels of Panha from a wrong National Park (Huu Lien N. R., Lang Son province), where both species do certainly not occur.
Therefore, the localities on the labels of Panha are with high probability wrong (see also Grego et al. 2014). The Cuc Phuong N. P. has obviously been changed by mistake with the Phong Nha – Ke Bang N. P., where both species have been collected by Ookubo and Hemmen.
Herewith, the type localities of Ptychauchenia panhai and Neniauchenia tonkinensis are corrected to Phong Nha – Ke Bang N. P., Quang Binh province.

Figs. 1-2. Garnieriinae with corrected type locality (phot. S. Hof).
Shell frontal, view on lunellar and body whorl dorsal. Shell height = H (mm).

Fig. 1. Ptychauchenia panhai H. Nordsieck, „Cuc Phuong N. P.“ = Phong Nha – Ke Bang N. P., holotype, SMF 331361, H 30.15.

Fig. 2. Neniauchenia tonkinensis H. Nordsieck, „Cuc Phuong N. P.“ = Phong Nha – Ke Bang N. P., holotype, SMF 331370, H 32.95.


Grego, J., Luong, H. V., Pham, S. V. & Szekeres, M. (2014): Vietnamese Clausiliidae (Gastropoda: Pulmonata): new taxa and novel distribution data. – Journal of Conchology, 41 (6): 749-757.

Nordsieck, H. (2010): New taxa of the subfamilies Neniinae and Garnieriinae (Gastropoda: Stylommatophora: Clausiliidae). – Archiv für Molluskenkunde, 139 (1): 45-69, 4 pls.

Nordsieck, H. (2011): Clausiliidae of Vietnam (Gastropoda, Stylommatophora) with the description of new taxa. – Archiv für Molluskenkunde, 140 (2): 149-173, 4 pls.