\nFormenkreis of septemplica<\/em>ta<\/i>:\u00a0Clausilia<\/i>\u00a0grohmanniana<\/i>\u00a0Rossm\u00e4ssler,\u00a0C<\/i>.\u00a0septemplicata\u00a0 <\/i>Philippi,\u00a0C<\/i>.\u00a0calcarae<\/i>\u00a0Philippi,\u00a0C<\/i>.\u00a0confinata<\/i>\u00a0=\u00a0scarificata<\/i>\u00a0L. Pfeiffer,\u00a0C<\/i>.\u00a0tiberii<\/i> A. Schmidt;
\nFormenkreis of\u00a0crassicostata<\/i>:\u00a0C<\/i>.\u00a0crassicostata<\/i>\u00a0L. Pfeiffer,\u00a0C<\/i>.\u00a0nobilis<\/i> L. Pfeiffer.
\nO. Boettger (1877: 33-34) named the groups of Schmidt (F. of\u00a0septemplicata<\/i>\u00a0=\u00a0Siciliaria<\/i>\u00a0Vest sensu stricto, F. of\u00a0crassicostata<\/i>\u00a0=\u00a0Trinacria<\/i>\u00a0O. Boettger). Later on, he (1879: 85-91) gave descriptions of several species taxa and listed eight species (C<\/i>.\u00a0leucophryne<\/i>\u00a0L. Pfeiffer added to those of Schmidt). His distribution data were in part based on wrong localities on the labels from Benoit and others.
\nBenoit (1882: 101-120) who had distributed most of the material in the collections of his correspondents listed 13 species, several of them regarded as varieties or synonyms by Boettger. Like those of Boettger, his distribution data were partially wrong.
\nWesterlund (1892: 47-48) published three of the numerous taxa names of Monterosato and gave a determination key for the species.
\nA. J. Wagner (1913, 1925) figured the genital organs of three\u00a0Siciliaria<\/i>\u00a0species from N. W. Sicily (S<\/i>.\u00a0septemplicata<\/i>: 1913: pl. 572, fig. 14; S<\/i>.\u00a0grohmanniana<\/i>: 1925: pl. 1, fig. 8,\u00a0S<\/i>.\u00a0calcarae<\/i>: pl. 3, fig. 25). He (1924: 123-125) gave a list of the species which is nearly identical with that of Boettger.
\nBrandt, after his collection trip in Sicily in 1949, prepared a manuscript with the description of 24 new taxa based on \u201especies\u201c of Monterosato or collected by himself. This manuscript remained unpublished; later on, only five of the species taxa were regarded by Brandt (1961: 6-9) as different enough to be described as new (among them two species:\u00a0S<\/i>.\u00a0ferrox<\/i>, with figure of genitalia fig. 1,\u00a0S<\/i>.\u00a0alcamoensis<\/i>). Besides, he (: 8) united S<\/i>.\u00a0leucophryne<\/i>\u00a0and\u00a0S<\/i>.\u00a0tiberii<\/i>\u00a0(and\u00a0S<\/i>.\u00a0eminens<\/i>\u00a0which later on was recognized as species) to one species under the name\u00a0S<\/i>.\u00a0leucophryne<\/i>, with which he also affiliated the newly described\u00a0S<\/i>.\u00a0riberothi<\/i> as subspecies. Many localities given by him cannot be identified or are obviously wrong.
\nAlzona (1971: 90-92) listed the same species as Boettger. Brandt’s new taxa were not considered, although his paper was cited.
\nNordsieck (1979: 259-260) listed also the same species as Boettger and (: 269) united his species groups (Siciliaria<\/i> s. str., Trinacria<\/i>\u00a0(preoccupied) =\u00a0Sicania<\/i>\u00a0Tomlin). Later on, he (1984: 202) described the new species\u00a0S<\/i>.\u00a0spezialensis<\/i> based on material collected by K. L. Pfeiffer.
\nManganelli et al. (1995: 25, 47-48), in the checklist of Italian pulmonate gastropods, enumerated 10 species taxa, proposed as such by Nordsieck (in litteris, S<\/i>.\u00a0ferrox<\/i>\u00a0and\u00a0S<\/i>.\u00a0spezialensis<\/i>\u00a0added to those of Nordsieck 1979). At the same time, Giusti et al. (1995: 327-330) evaded a distinction of species within the group which they gave the oldest name\u00a0S<\/i>.\u00a0septemplicata<\/i>. Therefore, they determined\u00a0S<\/i>.\u00a0calcarae<\/i> from Quaternary deposits of Malta as S<\/i>. cf.\u00a0septemplicata<\/i>.
\nNordsieck (2002: 30, 34) recognized\u00a0C<\/i>.\u00a0leucophryna<\/i>\u00a0and\u00a0C<\/i>.\u00a0eminens<\/i> as different species. He (29, 34) listed 12 species of Charpentieria<\/i>\u00a0(Siciliaria<\/i>) and classified them with three species groups, based on some shell characters regarded as informative: 1\u00a0nobilis<\/i>–calcarae<\/i>\u00a0group, 1a\u00a0nobilis<\/i>\u00a0subgroup (C<\/i>.\u00a0nobilis<\/i>,\u00a0C<\/i>.\u00a0spezialensis<\/i>,\u00a0C<\/i>.\u00a0crassicostata<\/i>,\u00a0C<\/i>.\u00a0eminens<\/i>), 1b\u00a0calcarae<\/i> subgroup (C<\/i>.\u00a0calcarae<\/i>,\u00a0C<\/i>.\u00a0tiberii<\/i>,\u00a0C<\/i>.\u00a0leucophryna<\/i>), 2\u00a0grohmanniana<\/i>\u00a0group (C<\/i>.\u00a0grohmanniana<\/i>,\u00a0C<\/i>.\u00a0septemplicata<\/i>,\u00a0C<\/i>.\u00a0scarificata<\/i>). Because of their ambiguous character combinations,\u00a0C<\/i>.\u00a0ferrox<\/i>\u00a0and\u00a0C<\/i>.\u00a0riberothi<\/i>\u00a0were not included in one of these groups. Several forms assumed to have originated by interspecies hybridization (: 34) were added.
\nBeckmann (2004: 188, maps 1 and 2) published detailed informations on the distribution of the species and described (: 188-190) two new subspecies, both assumed to be hybrid forms.
\nReitano et al. (2007: 326-328) gave further informations on the distribution of\u00a0S<\/i>.\u00a0ferrox<\/i>.
\nLiberto et al. (2015) published a detailed description of S<\/em>. scarificata<\/em>.
\nLiberto et al. (2016) gave further informations on morphology and distribution of\u00a0 S<\/em>. calcarae<\/em> and (: 371) described a new subspecies of that species, S<\/em>. c<\/em>. orlandoi.
\n<\/em>De Mattia et al. (2021) published a comprehensive study of the Siciliaria<\/em> species from the region under consideration, with focus on the genital morphology, together with a DNA analysis. This paper is based only on the material collected by De Mattia and Macor; the material which has been treated in this article has not been considered by the authors<\/p>\n
II. System<\/h4>\n
The material on which the following system is based comprises about 230 samples of Siciliaria<\/i>\u00a0(S<\/i>.) species from the collection of the Forschungsinstitut Senckenberg (abbreviation: SMF), containing the types of several species taxa, and 80 samples of my own collection, deposited in the Staatliches Museum f\u00fcr Naturkunde Stuttgart (abbreviation: SMNS) and the Forschungsinstitut Senckenberg. In the Senckenberg collection material collected by Kobelt, K. L. Pfeiffer, Brandt and Beckmann is kept, in my own collection an important part of the Brandt collection (SMNS) and material collected by C. and J. Hemmen in 2009 (SMF). 80 samples of all taxa have been subjected to a more thorough morphological examination.
\nIn collections from the nineteenth century the localities given on the labels have often turned out to be wrong. In this article only locality data given by collectors, who in my opinion are reliable, are used. In any case, I had to rely on those data, because until now I had not the opportunity to collect in this region.
\nThe examination of the genital morphology of several Siciliaria<\/i> species (see website article on Genital studies) had the result that the genitalia of all species are similar.<\/p>\n1. Diagnoses and distribution<\/h5>\n
In the following the diagnoses of the species of Siciliaria<\/i>\u00a0from N. W. Sicily and their distribution, as far as known from the examined material, are given. In notes, geographic forms which could have subspecific rank are characterized. Among them, there are several forms which have probably originated by interspecies hybridization (see chapter 2.). Figs. 1-7.\u00a0Siciliaria\u00a0<\/i>spp. <\/p>\n Siciliaria<\/i>\u00a0Vest 1867 For the subdivision of\u00a0Siciliaria<\/i>\u00a0from N. W. Sicily see website art. on DNA studies.<\/p>\n S.\u00a0<\/i>(Siciliaria<\/i>)<\/p>\n Nomenclatural note:\u00a0Clausilia<\/i>\u00a0decollata<\/i>\u00a0and\u00a0C<\/i>.\u00a0deshayesii<\/i>\u00a0Potiez & Michaud 1838 are unidentified species. The first may be\u00a0S<\/i>.\u00a0grohmanniana<\/i>, the second\u00a0S<\/i>.\u00a0septemplicata<\/i>\u00a0or\u00a0S<\/i>.\u00a0calcarae<\/i>. Three \u201especies\u201c of Benoit (1882) are also unidentified: C<\/i>.\u00a0trinaclina<\/i>\u00a0(misprint for\u00a0trinacrina<\/i>) (=\u00a0S<\/i>.\u00a0calcarae<\/i>?),\u00a0C<\/i>.\u00a0rubra<\/i>\u00a0(=\u00a0S<\/i>.\u00a0septemplicata<\/i>?) and\u00a0C<\/i>.\u00a0subdiaphana<\/i>\u00a0(=\u00a0S<\/i>.\u00a0grohmanniana<\/i>?). The variety names published by Monterosato, in fact infrasubspecific, are not considered in this article.<\/p>\n S<\/i>.\u00a0grohmanniana<\/i>\u00a0(Rossm\u00e4ssler 1836) Diagnosis: Shell mostly decollated; whorls ribbed; dorsal keel indistinct or missing; inferior lamella very high; two anterior upper palatal plicae present, upper one mostly separated from upper palatal plica; palatal edge of clausilium plate distally receding, plate gutter-like narrowed, palatal edge against distal end bent up and more or less pointed.<\/p>\n Distribution: Monte Pellegrino, Monte Gallo.<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0grohmanniana<\/i>\u00a0var.\u00a0minor<\/i>\u00a0A. Schmidt 1868 [non Rossm\u00e4ssler],\u00a0C<\/i>.\u00a0grohmanniana<\/i>\u00a0var.\u00a0panormitana<\/i>\u00a0Paulucci 1878.<\/p>\n Intraspecific forms: S<\/i>.\u00a0septemplicata<\/i>\u00a0(Philippi 1836) Diagnosis: Shell mostly not decollated; whorls rib-striated, with sutural papillae; dorsal keel indistinct or missing; inferior lamella very high; two anterior upper palatal plicae present, upper one mostly separated from upper palatal plica; palatal edge of clausilium plate distally receding, plate gutter-like narrowed, palatal edge against distal end bent up and more or less pointed.<\/p>\n Distribution: From Cinisi in the west and Monte Gallo in the north to Piana degli Albanesi in the south and Monte Catalfano in the east.<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0sericina<\/i>\u00a0Rossm\u00e4ssler 1836,\u00a0C<\/i>.\u00a0septemplicata<\/i>\u00a0var.\u00a0prasina<\/i> A. Schmidt 1868, C<\/em>. allyphanta<\/em> Westerlund 1894. The name C<\/i>.\u00a0prasina<\/i>\u00a0is a copying-mistake of\u00a0C<\/i>.\u00a0proxima<\/i>, caused by illegible labels of Benoit.<\/p>\n Intraspecific forms: S<\/i>.\u00a0leucophryna<\/i>\u00a0(L. Pfeiffer 1862)<\/p>\n Diagnosis: Shell decollated; whorls ribbed; dorsal keel mostly indistinct or missing; inferior lamella moderately high; anterior upper palatal plica present, separated from or connected with upper palatal plica; palatal edge of clausilium plate distally slightly or not receding, palatal edge distally somewhat or not bent up.<\/p>\n Distribution: Isola delle Femmine and Grotta Conza to Monte Gallo.<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0leucophryne<\/i>\u00a0var.\u00a0laudabilis<\/i>\u00a0O. Boettger 1879,\u00a0C<\/i>.\u00a0confinata<\/i>\u00a0var.\u00a0merens<\/i>\u00a0Westerlund 1892.<\/p>\n Intraspecific forms: S<\/i>.\u00a0calcarae<\/i>\u00a0(Philippi 1844) Diagnosis: Shell not decollated; whorls rib-striated, with sutural papillae; dorsal keel indistinct or missing; inferior lamella moderately high or low; anterior upper palatal plica present, mostly separated from upper palatal plica, rarely lower anterior upper palatal plica present; palatal edge of clausilium plate distally not receding, palatal edge distally more or less strongly bent up.<\/p>\n Distribution: Island of Favignana, whole mainland of N. W. Sicily from Erice in the west to Montagna Grande in the east; in the south to Calatafimi and Ficuzza region.<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0adelinae<\/i>\u00a0K\u00fcster 1860,\u00a0C<\/i>.\u00a0brugnoneana<\/i>\u00a0Pini 1884,\u00a0C<\/i>.\u00a0calcarae<\/i>\u00a0var.\u00a0nodosa<\/i>\u00a0Westerlund 1892.<\/p>\n Intraspecific forms: S<\/i>.\u00a0tiberii<\/i>\u00a0(A. Schmidt 1868) Diagnosis: Shell not decollated or decollated; whorls ribbed; dorsal keel of different prominence; inferior lamella moderately high or low; anterior upper palatal plica present, mostly separated from upper palatal plica, rarely lower anterior upper palatal plica present; palatal edge of clausilium plate distally not receding, palatal edge distally more or less strongly bent up.<\/p>\n Distribution: From north of Cinisi in the north to San Cipirello in the south, Monte San Calogero?<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0tiberiana<\/i>\u00a0Paulucci 1878.<\/p>\n Intraspecific forms: S.\u00a0ferrox<\/i>\u00a0Brandt 1961<\/p>\n Diagnosis: Shell mostly not decollated; whorls ribbed, sutural papillae recognizable; dorsal keel distinct or indistinct; inferior lamella moderately high or high; two anterior upper palatal plicae present, upper one mostly separated from upper palatal plica; palatal edge of clausilium plate distally less or not receding, palatal edge distally not bent up.<\/p>\n Distribution: From Altavilla Milicia to Termini Imerese.<\/p>\n Synonyms: C.\u00a0confinata<\/i>\u00a0var.\u00a0commeata<\/i>\u00a0Westerlund 1892 could be an older synonym, but the description is too poor (lower anterior upper palatal plica not mentioned) and types are not available (Reitano et al. 2007: 328).<\/p>\n S<\/i>. (Sicania<\/i>) Tomlin<\/p>\n S<\/i>.\u00a0nobilis<\/i>\u00a0(L. Pfeiffer 1848) Diagnosis: Shell decollated; with well-developed white surface layer; whorls mostly weakly rib-striated, with sutural papillae; dorsal keel missing; inferior lamella moderately high or high; anterior upper palatal plica present, mostly separated from upper palatal plica, or missing, basalis tending to be shortened, posterior lower palatal plica more or less reduced; palatal edge of clausilium plate distally not receding, palatal edge distally somewhat or not bent up.<\/p>\n Distribution: San Vito peninsula from Monte Monaco and Monte Acci, Monte Cofano.<\/p>\n Synonyms:\u00a0C<\/i>.\u00a0cantrainei<\/i> Deshayes 1851.<\/p>\n Intraspecific forms: S<\/i>.\u00a0spezialensis<\/i>\u00a0H. Nordsieck 1984<\/p>\n Diagnosis: Shell decollated; with well-developed white surface layer; whorls ribbed or weakly rib-striated, with sutural papillae; dorsal keel missing; inferior lamella mostly high; anterior upper palatal plica missing, basalis missing, posterior lower palatal plica connected with lunella; clausilium plate fit in the frame of lunella and posterior lower palatal plica; palatal edge distally not bent up.<\/p>\n Distribution: San Vito peninsula from Macari and Castelluzzo to Monte Acci and Monte Speziale.<\/p>\n Intraspecific forms: S<\/i>.\u00a0eminens<\/i>\u00a0(A. Schmidt 1868)<\/p>\n Diagnosis: Shell decollated; whorls ribbed; dorsal keel indistinct or missing; inferior lamella mostly high; anterior upper palatal plica present, mostly separated from upper palatal plica, basalis tending to be shortened; palatal edge of clausilium plate distally not receding, palatal edge distally somewhat or not bent up.<\/p>\n Distribution: From Custonaci in the west to Castellammare in the east, environs of Calatafimi, M. Bonifato near Alcamo.<\/p>\n Intraspecific forms: S<\/i>.\u00a0crassicostata<\/i>\u00a0(L. Pfeiffer 1856) Diagnosis: Shell decollated; whorls ribbed, in part percostate; dorsal keel distinct or indistinct; inferior lamella moderately high or high; anterior upper palatal plica present, separated from upper palatal plica, in part missing, basalis tending to be shortened; palatal edge of clausilium plate distally not receding, palatal edge distally somewhat or not bent up.<\/p>\n Distribution:: Monte Cofano, (probably introduced on) island of Favignana.<\/p>\n Siciliaria\u00a0<\/i>(Mauritanica<\/em>) O. Boettger 1879<\/p>\n S<\/i>.\u00a0scarificata<\/i>\u00a0(L. Pfeiffer 1856)<\/p>\n Diagnosis: Shell mostly decollated; whorls ribbed, sutural papillae recognizable; dorsal keel indistinct ore missing; inferior lamella high; anterior upper palatal plica present, separated from or connected with upper palatal plica; palatal edge of clausilium plate distally receding, palatal edge distally somewhat or not bent up.<\/p>\n Distribution: Island of Marettimo.<\/p>\n Synonyms: The names\u00a0C<\/i>.\u00a0scarificata\u00a0<\/i>and\u00a0C<\/i>.\u00a0sacrificata<\/i>\u00a0are copying-mistakes of\u00a0C<\/i>.\u00a0confinata<\/i>, caused by illegible labels of Benoit.<\/p>\n The forms of Siciliaria<\/i>, which are possibly hybrid ones, are enumerated in the following. They are forms of well-defined species restricted to certain localities or regions differing from the nominotypical forms by showing characters of another species. This other species is clearly different from that species and is distributed in the same region. The possible hybrid forms occur in the neighbourhood of or are even surrounded by populations of the nominotypical form of the species to which they belong. They could be the products of gene flow from the other species restricted to a limited locality or region.
\nThe following shell characters have turned out to be informative for species diagnoses (see figs. 1-7):
\n1. Decollation: Shell entire \/ decollated.
\n2. Sculpture: Whorls rib-striated, with sutural papillae \/ weakly ribbed, papillae still recognizable \/ strongly ribbed.
\n3. Cervix of body whorl: Only with basal keel, without dorsal keel \/ with more or less pronounced dorsal keel.
\n4. Inferior lamella: Lamella more or less protruding into the lumen of the body whorl, described as: lamella high \/ moderately high \/ low.
\n5.<\/b>\u00a0Anterior upper palatal plica: Only one plica present \/ two plicae present, upper and lower one \/ plica missing.
\n6. Clausilium plate: Palatal edge of plate distally receding, thus distal part narrowed, forming a gutter (figs. 1-2) \/ not receding, palatal and columellar edges of plate nearly parallel (figs. 3-7). Palatal edge distally much bent up (figs. 3-4) \/ bent up (figs. 1-2) \/ somewhat or not bent up (figs. 5-7).
\nThe distribution of the species concerned is restricted to N. W. Sicily: Egadi islands, Sicilian mainland from Trapani in the west to the western boundary of Madonie in the east, in the south delimited by the imaginary line Trapani\u2013Calatafimi\u2013San Cipirello\u2013Ficuzza\u2013Monte San Calogero.
\nIn a larger part of N. W. Sicily two species pairs with identical clausilium plate, but different appearance (shell entire, nearly smooth and papillate \/ shell decollated, ribbed) occur (figs. 1-4). The smooth papillate species is adapted to more shady habitats, while the ribbed one lives in more open rock habitats. Decollation is possibly an adaptation to a life under stones. One of these species pairs,\u00a0S<\/i>.\u00a0septemplicata<\/i>\u00a0\/\u00a0S<\/i>.\u00a0grohmanniana<\/i>\u00a0(figs. 1-2), is distributed in the east, the other one,\u00a0S<\/i>.\u00a0calcarae<\/i>\u00a0\/\u00a0S<\/i>.\u00a0tiberii<\/i>\u00a0(figs. 3-4), in a larger part of the region,\u00a0S<\/i>.\u00a0calcarae<\/i>\u00a0in the whole region,\u00a0S<\/i>.\u00a0tiberii<\/i> only in the central and eastern part. The range restriction of the latter is probably caused by the occurrence of other rock-dwelling species, those of Sicania<\/em>, in the west.
\nOn the San Vito peninsula, there is a species pair of Sicania<\/em>, which lives in open rock habitats, S<\/i>.\u00a0nobilis<\/i>\u00a0\/\u00a0S<\/i>.\u00a0spezialensis<\/i>. Both are adapted to this habitat by a well-developed white surface layer of the shell (fig. 5), looking like\u00a0Albinaria<\/i>\u00a0species, both are decollated.\u00a0S<\/i>.\u00a0spezialensis<\/i> exhibits a special closing apparatus (clausilium plate fit into a frame of palatal plicae, anterior palatal plicae missing), which may better protect it from desiccation. On one mountain of this peninsula, Monte Cofano, a species with percostate shell, S<\/i>.\u00a0crassicostata<\/i>\u00a0(fig. 6), occurs. This sculpture, which is found also in certain\u00a0Albinaria<\/i>\u00a0species and other rock-dwelling clausiliids of meridional Europe, may be another adaptation to a life in open rock habitats.
\nThe occurrences of\u00a0Siciliaria<\/i> species on the Egadi islands are of special interest. On Marettimo, only S<\/i>.\u00a0scarificata<\/i>\u00a0occurs. On Favignana, originally only\u00a0S<\/i>.\u00a0calcarae<\/i> occurred. S.\u00a0<\/em>crassicostata<\/i>, which has been collected by several authors on the island (e. g., by Brandt on garden walls, fig. 6) has obviously been introduced (by man?) from Monte Cofano nearby.<\/p>\n
\n<\/strong>Frontal view and body whorl dorsal view. Actual shell height = H (mm). Phot. S. Hof.<\/p>\n![\"\"](\"https:\/\/hnords.de\/wordpress\/wp-content\/uploads\/2021\/05\/tafelsiciliaria1a.jpg\")
<\/a><\/p>\nFig. 1.\u00a0S<\/i>. s<\/em>. septemplicata<\/i>, Passo di Rosetto near Palermo, ex SMF 67942, H = 19.5, shell with clausilium plate H = 19.0.
\nFig. 2.\u00a0S<\/i>.\u00a0grohmanniana<\/i>, Monte Pellegrino near Palermo, ex SMF 67528, H = 19.3, shell with clausilium plate H = 18.1.
\nFig. 3.\u00a0S<\/i>. c<\/em>. calcarae<\/i>, Sferracavallo, ex SMF 67925, H = 19.0.
\nFig. 4.\u00a0S<\/i>.\u00a0tiberii riberothi<\/i>, Monte della Fiera, ex SMF 333547, H = 17.1.<\/h5>\n<\/a><\/h5>\n<\/h5>\n
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Fig. 5.\u00a0S<\/i>.\u00a0nobilis<\/i>, Capo San Vito, ex SMF 96476, H = 20.9.
\nFig. 6.\u00a0S<\/i>.\u00a0crassicostata<\/i>, Island of Favignana, ex SMF 265742, H = 18.9.
\nFig. 7.\u00a0S<\/i>. tiberii<\/em> riberothi<\/i>, \u201eMonte della Terrazza\u201c, ex SMF 232084, H = 20.2.<\/h5>\n
\n<\/i><\/p>\n
\nFig. 2<\/p>\n
\nPriola form =\u00a0g<\/em>. addaurae<\/em> De Mattia et al. 2021, Priola near Palermo: Shell not decollated; whorls more weakly ribbed.<\/p>\n
\nFig. 1<\/p>\n
\nParco form, Parco near Palermo: Inferior lamella in part lower; lower anterior upper palatal plica missing; palatal edge of clausilium plate less or not bent up.
\ns<\/i>.\u00a0alcamoensis<\/i> Brandt 1961, \u201eAlcamo\u201c: Cervix more strongly sculptured; dorsal keel more pronounced; inferior lamella in part lower; lower anterior upper palatal plica more or less reduced; clausilium plate distally less narrowed, palatal edge less upbent. s<\/i>.\u00a0hemmeni<\/i> (Beckmann 2004), Piano Margi near Cinisi, is considered as synonym.<\/p>\n
\nSferracavallo form, Sferracavallo: Inferior lamella lower; palatal edge of clausilium plate more strongy bent up.<\/p>\n
\nFig. 3<\/p>\n
\nc<\/i>.\u00a0belliemii<\/i>\u00a0Brandt 1961, Monte Belliemi near Partinico: Whorls ribbed.
\nSagana form =\u00a0c<\/em>. borgettensis<\/em> De Mattia et al. 2021, Sagana near Montelepre: Inferior lamella higher; lower anterior upper palatal plica more or less developed; palatal edge of clausilium plate less strongly bent up.
\nc<\/em>. cruenta<\/em> De Mattia et al. 2021, Monte Gibilmesi near Montelepre: Like c<\/em>. borgettensis<\/em>, but darker coloured.
\nc. orlandoi <\/i>Liberto, Reitano, Giglio, Colomba & Sparacio 2016, region of Ficuzza: Dark brown; no further anterior upper palatal plica present; clausilium plate like in S<\/i>.\u00a0septemplicata<\/i>, but distally less narrowed.<\/p>\n
\nFigs. 4, 7, 10<\/p>\n
\nSouthern form from Monte Jato and Monte Kumeta = t<\/em>. tiberii<\/em>, Monte San Calogero?: Shell not or only in part decollated; dorsal keel less strong.
\nThere is no doubt that\u00a0Clausilia tiberii<\/i> A. Schmidt (1868: 42-43) belongs to the southern form. Originals from Benoit (SMF 67544, 96458, see also Benoit 1859: pl. 6, fig.7) do not differ from the form from Monte Jato and Monte Kumeta (also from Monte della Scala fig. 10).
\nNorthern form from Monte Palmeto and Monte della Fiera = t<\/em>. riberothi<\/em>: Shell decollated; dorsal keel strong.
\nS<\/i>.\u00a0riberothi\u00a0<\/i>Brandt (fig. 7) is similar to the northern form (see chapter 3.), differing mainly by the clausilium plate with palatal edge distally not bent up. Therefore, the name riberothi<\/em> is used for the northern form.
\nArmetta form =\u00a0t.<\/em> armettensis<\/em> De Mattia et al. 2021, Grotta Armetta near Carini: Shell decollated; more densely ribbed; dorsal keel weak; inferior lamella higher.
\nt<\/i>.\u00a0scalettensis<\/i>\u00a0(Beckmann 2004), Port la Scaletti near Cinisi: Whorls rib-striated; inferior lamella higher; palatal edge of clausilium plate somewhat or not bent up.<\/p>\n
\nFig. 5<\/p>\n
\nType form from \u201eRedicofano\u201c = probably San Vito: Anterior upper palatal plica mostly present.
\nForm from remaining species range: Anterior upper palatal plica mostly missing.
\nepisoma <\/i>form, coast near S. Vito: Shell smaller; whorls more distinctly rib-striated; anterior upper palatal plica missing.<\/p>\n
\nType form is ribbed.
\nForm from remaining species range: Whorls weakly rib-striated.<\/p>\n
\nCustonaci form: Whorls more widely ribbed, transitional to S.<\/em> crassicostata<\/em>.<\/p>\n
\nFig. 6<\/p>\n2. Interspecies hybridization<\/h5>\n
\nThese forms are the following:
\n1. S.\u00a0grohmanniana addaurae<\/i>\u00a0(fig. 8), with characters of S<\/i>.\u00a0septemplicata<\/i>, occurring in the environs of Palermo, where both species are distributed.
\n2.\u00a0S<\/i>.\u00a0septemplicata<\/i>, Parco form, with characters of\u00a0S<\/i>.\u00a0calcarae<\/i>, from a locality near Palermo in a region, where both species have been traced in localities nearby.
\n3.\u00a0S<\/i>.\u00a0septemplicata<\/i>\u00a0alcamoensis\u00a0<\/i>(fig. 9) =\u00a0S<\/i>.\u00a0s<\/i>.\u00a0hemmeni<\/i>, with characters of\u00a0S<\/i>.\u00a0tiberii<\/i>, the latter occurring near Cinisi in a region, where both species are distributed. The locality Alcamo for\u00a0S<\/i>.\u00a0s<\/i>.\u00a0alcamoensis<\/i>\u00a0is probably wrong, also because neither\u00a0S<\/i>.\u00a0septemplicata<\/i>\u00a0nor\u00a0S<\/i>.\u00a0tiberii<\/i>\u00a0have been found in that region.
\n4. S<\/i>.\u00a0calcarae<\/i>\u00a0belliemii<\/i>\u00a0from Monte Belliemi, with the ribbing of\u00a0S<\/i>.\u00a0tiberii<\/i>, which occurs in the Monte della Fiera nearby.
\n5. S<\/i>.\u00a0t.<\/i>\u00a0tiberii<\/i>, type form from Monte della Scala (fig. 10), with characters of